Tsukuyomaru

was a bat demon and the father of Shiori. He fell in love with a human woman and they had a hanyō daughter together, Shiori. His father considered him weak for doing so. Father and son would later have an argument about this, with Tsukuyomaru having decided to leave his demon clan for the human village where his wife and daughter lived. He would also abandon the hereditary duty of creating a barrier with the red orb. His father's hatred then came to a head and he killed Tsukuyomaru without any remorse, planning to use Shiori instead to care for the red orb.

Name Meaning
Tsukuyomaru literally means "moonlit night circle".

Trivia
When Tsukuyomaru folds his wings over his body, his wings look like a cape.